The government has denied as "baseless" accusations it was behind the killing. However, a police statement noted that one of the suspects is related to someone allegedly killed by Rugambage during the 1994 genocide.
Rugambage, who is survived by his wife and a child, was acquitted of genocide crimes by a local "gacaca" court in 2006.
According to the BBC, Human rights groups have accused President Paul Kagame of intimidating the media and the opposition ahead of elections due to take place in August.
A police statement quoted by the pro-government New Times newspaper said the pistol used to shoot Rugambage has been recovered.
The authorities recently suspended the Umuvugizi paper, prompting it to start publishing online instead.
The report quoted the editor Jean Bosco Gasasira, who fled to Uganda in April after his paper was suspended, to have said Kigali had masterminded the assassination of Mr Rugambage who died in hospital after the shooting.
"I'm 100% sure it was the office of the national security services which shot him dead," he told the Voice of America.

With effect from 1st October, 2007Tanzania National Parks started using electronic payment systems at its revenue collection centres in parallel run with the old system. Phase I of this system has covered Serengeti, Lake Manyara, Tarangire, Kilimanjaro and Arusha National Parks through CRDB and EXIM Banks who designed the systems.













President of the Republic of Tanzania, His Excellence, Dr. Jakaya Kikwete, said there are several investment opportunities in Tanzania which awaits eligible businessmen to invest.
All 45 members of the group of celebrities and professionals who are in the campaign on the Global Clean Water Crisis – Summit on the Summit made it to the top of the African highest Mountain – Kilimanjaro.
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